The Lion

There are at least fifty species of cats, belonging to the family of Felidae (mammalian order of carnivores). They are built on much the same plan, but for convenience are divided into the big and the not so big cats. Those designated as the big cats (genus Panthera) include the lion, tiger, leopard, jaguar, clouded leopard and snow leopard. The cheetah, however, is a somewhat different kind of feline. The cat known as the American, Mexican, or mountain lion is a New World member of the genus Felis (see puma).

The lion (Leo, or Panthera, leo): large, powerfully built cat of the family Felidae, and the second largest of the big cats (after the tiger). The proverbial "king of beasts," the lion has been, since earliest times, one of the best known of wild animals. It is now found mainly in parts of Africa south of the Sahara. A few hundred lions, constituting an Asiatic race, live under strict protection in the Gir Forest National Park in Gujarat state, India. The preferred habitats of lions are grassy plains and open savannah.

Physics

The lion is a well-muscled cat with a long body, short legs, and large head. It varies considerably between the sexes in both size and appearance. A full-grown male is about 1.8-2.1 m (6-7 feet) long, excluding the 1-metre (3-foot) long tail; stands about 1.2 m (4 feet) high at the shoulder; and weighs 170-230 kg (370-500 pounds). The female, or lioness, is smaller, with a body length of 1.5 m (5 feet), a shoulder height of 0.9-1 m (3-3.5 feet), and a weight of 120-180 kg (265-390 pounds). The lion's coat is short and varies in colour from buff yellow, orange-brown, or silvery grey to dark brown, with a tuft on the tail tip that is usually darker than the rest of the coat. Lionesses are more consistently tawny or sandy in colour. The male's outstanding characteristic is his mane, which varies in different individuals. It may be entirely lacking; it may fringe the face; or it may be full and shaggy, covering the back of the head, neck, and shoulders and continuing onto the throat and chest to join a fringe along the belly. The mane and fringe of some lions are very dark, almost black; that gives the animals a majestic appearance.

Big cats are outstanding among the carnivores, or flesh-eating animals, for their unsurpassed combination of muscular power and grace of movement. These magnificent creatures generally catch their prey by clever stalking or ambush. They must be on the alert and tensed to make the kill; at the same time they must remain quiet enough not to frighten the intended victim. Sometimes they pursue game that they can neither smell nor see; in such cases they rely upon such clues as footmarks.

The cat's body has great elasticity. Because the vertebrae of the spinal column are held together by muscles rather than by ligaments, as in humans, the cat can elongate or contract its back, curve it upward, or oscillate it along the vertebral line. The construction of the shoulder joints permits the cat to turn its foreleg in almost any direction. Cats are powerfully built animals and are so well co-ordinated that they almost invariably land on their feet if they fall or are dropped.

The long and lithe bodies of the felines are wonderfully specialised for their hunting activities. A good deal of the weight is carried on the forelimbs. Strong back and hindleg muscles enable the animals to make sudden leaping movements as they pounce upon the prey. The cats are not as sturdy runners as the dogs; instead of steadily pursuing the prey for minutes at a time, they make rapid dash followed by a sudden leap. Cats are digitigrade; that is, they walk on their toes; there are five toes on each front and four on each hind foot. Unlike the dog and horse, the cat walks or runs by moving first the front and back legs on one side, then the front and back legs on the other side; only the camel and the giraffe move in a similar way. The smaller cats are adept at climbing trees; the large felines live mostly on the ground.

Head and teeth

A cat's head is large, while the face is relatively short and rounded. There is a bony crest at the top of the skull for the attachment of the powerful temporal muscles that manipulate the lower jaw. The animal has highly specialised teeth. The front teeth, or incisors, serve to bite off flesh; the fangs, or canine teeth, are efficient daggers for piercing and slashing. Finally, there are modified teeth, the carnassials, which shear flesh and cut though sinews.



The cat's teeth are adapted to three functions: stabbing (canines), anchoring (canines), and cutting (molars). Cats have no flat-crowned crushing teeth and therefore cannot chew their food; instead, they cut it up. Except for the canines and molars, the cat's teeth are more or less non-functional; most of the cheek teeth do not even meet when the mouth is closed.



The carnassials, which are shaped like cutting blades, are a combination of the last premolar of the upper jaw and the first molar of the lower jaw. As the jaws close, the upper blade moves past the lower one; the action suggests that of a pair of scissors.

Tsavo tooth

Almost 3/4 of this 10 cm (4 inch) long lion tooth is its root, which holds the tooth in place as the animal bites and subdues its prey.The dental formula in all cats, for either side of both upper and lower jaws, is incisors 3/3, canines 1/1, premolars 3/2, and molars 1/1. The total number of teeth is 16 in the upper jaw and 14 in the lower. Primary, or milk, teeth number 24; these are replaced by the permanent teeth at about five months. Each half of the jaw is hinged to the skull by a transverse roller that fits tightly into a trough on the underside of the skull, making grinding movements impossible even if the cat had teeth suitable for grinding.

Claws

The toes are armed with long, curved and sharp claws, which can be drawn back, or retracted, when not in use. The claws are primarily utilised for seizing prey. There is a remarkable mechanism for retracting the cat's claws when they are not in use. The claw is retracted or extended by pivoting the end bone of the toe, which bears the claw, over the tip of the next be.

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The action that unsheathes the claws also spreads the toes widely, making the foot more than twice as broad as it normally is and converting it into a truly formidable weapon. This claw-sheathing mechanism is present in all species of the cat family except the cheetah. Although there are no nerve endings in the nail itself, blood capillaries are present in the inner part.

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Senses

Cats are generally nocturnal in habit. The retina of the cat's eye is made extra sensitive to light by a layer of guanine, which causes the eye to shine at night in a strong light. The eyes themselves, large with pupils that expand or contract to mere slits according to the density of light, do not distinguish colours clearly, but they are perfected for gathering the maximum amount of reflected light available to them at night. In open country, especially, cats use their eyes to detect quarry. Cats have a third eyelid, or nictitating membrane, commonly called the haw. Its appearance is used frequently as an indicator of the cat's general state of health.

The cat's sense of smell, particularly well developed in the adult, is crucial to its evaluation of food, so that a cat whose nasal passages become clogged as a result of illness may appear to lose its appetite completely. Cats can distinguish the odour of nitrogenous substances (e.g., fish) especially keenly.

The sense of touch is acute in cats. The eyebrows, whiskers, hairs of the cheek, and fine tufts of hair on the ears are all extremely sensitive to vibratory stimulation. The functions of the whiskers (vibrissae) are only partially understood; however, it is known that, if they are cut off, the cat is temporarily incapacitated. The toes and paws, as well as the tip of the nose, are also very sensitive to touch.

On the tongue there are backward curved prickles that provide a rough surface for rasping particles of meat from a bone and has the appearance and feel of a coarse file; the spines help the cat to lap up liquids and also to groom itself. The disposition to cleanliness is well established in cats, and they groom themselves at length, especially after meals.

Cats also have an acute sense of hearing. Their ears contain almost 30 muscles (compared with 6 in humans); as a result, they can turn them many times more quickly in the direction of a sound than can a dog. The ears of cats, although receptive to ultrasonic frequencies up to 25,000 vibrations per second, are slightly inferior to those of dogs, which register 35,000 vibrations per second.

A cat's brain is large and well developed, making the animal extremely alert and controlling its highly co-ordinated actions.


Nourishment

Cats prey on any animal that they are quick enough to catch and strong enough to subdue. Lions prey on a large number of animals ranging in size from gazelles and baboons upward to buffalo and hippopotamuses, but they prefer to hunt such medium- to large-sized hoofed animals as wildebeest, zebra, and impala and other antelopes. Lions readily eat any meat they can find, including carrion and fresh kills that they scavenge from hyenas through the use of force or intimidation. Although felines are almost exclusively meat eaters, the lion and leopard are known to feed on vegetable matter, principally fallen fruit. The lion will also eat rotten wood and ashes, when the animals it preys on become scarce.

It is the lionesses of a pride that do most of the hunting. When hunting, the cats pay no attention to the wind's direction, which can carry their scent to their prey, and they also tire after running only short distances; a high proportion of their hunts thus end in failure. Normally a male is circling around the herd of prey animals, while the lionesses patiently wait in ambush, using every bit of available cover.

The hunters keep in communication by emitting grunts every so often. The male gets upwind of the quarry so that his scent is carried to them. He singles out several individuals, mostly the weakest animals and tries to cut them off from the rest of the herd, meanwhile driving the prey toward the hidden lionesses. Once a victim us within range, there is little hope of its escape. The lioness makes a short, high-speed charge, up to fifty miles an hour, and usually leaps at the shoulder area and uses the powerful forepaw with claws extended to wrench back the victim's head violently, as the teeth sink in the back of the neck, until the animal is strangled. The kill is swift and clean. Other members of the pride quickly crowd around to feed on the kill, with males obtaining the most meat in the resulting scuffles and the smaller cubs acquiring little or none. The carcass is disembowelled, the heart and liver being especially prized. These animals do not crush the bones, nor pick them completely clean. This leaves enough to feed for the jackals, hyenas and vultures that always follow in the train of the great hunters.

Of course, the male is perfectly capable of making the kill, but his function is usually that of co-ordinating the movements of its troop. Another, less successful strategy is to approach a herd from opposite directions and then closing in to try for a kill in the resulting panic. The hunting group stands a better chance when they concentrate on one victim.

Lions and lionesses typically gorge themselves on a kill and then rest for several days in its vicinity. After consuming more than 34 kg (75 pounds) of meat at a single meal, an adult male can rest for a week before resuming the hunt. If prey is abundant, both sexes typically spend 20 hours a day resting, sleeping, or sitting and only hunt for 2 or 3 hours a day.


Behaviour

Lions are unique among cats in that they live in a group, or pride. A pride consists of several generations of lionesses, all of whom are related, their cubs, and one or two adult male lions that defend the pride's territory and mate with the females. The adult males are outsiders who may hold the pride for a few months to several years, depending on their ability to defend it against other outsider males. Candidates for the job are must be in excellent physical condition. Normally, fighting among lions is a rare occurrence, though frequently a lion who is a stranger to the vicinity is attacked; male lions may battle fiercely, with sometimes fatal results, for possession of a sexually receptive lioness. When the troop leader shows signs of a decline of power and speed, the younger, more vigorous males quickly drive him out from the group. The former leader then lives alone, depending more and more on smaller animals and food, until he dies. His remains are as often as not devoured by the members of his former troop. The fate of the old lioness is not so stern, for she is allowed to follow the group and feed on the kills of the others.

A pride may have as few as 4 or as many as 37 members, but about 15 is the average size. Each pride has a well-defined territory; where prey is abundant, its area may be as small as 20 square km (8 square miles), but if game is sparse, the territory may be up to 400 square km (150 square miles) in area. Male cubs are expelled from the pride at about 3 years of age and become nomads until they are old enough (at age 5) to try and take over another pride. Many adult males remain nomads for life, however. Some female cubs remain within the pride when they attain sexual maturity, but others are forced out and join other prides. The members of a pride typically spend the day in several scattered groups, though these may unite to hunt or to share in a kill.

Lions proclaim their territory by roaring and by scent marking. The lion's well-known roar is generally uttered in the evening before a night's hunting and again before getting up at dawn. Males also proclaim their presence by urinating on bushes, trees, or simply on the ground, leaving pungent scent markings in the process. Defecation and rubbing against bushes leave other scent markings.

Both sexes are polygamous and breed throughout the year, but females are usually restricted to the one or two adult males of a pride for their breeding partners. In captivity lions often breed every year, but in the wild they usually breed no more than once in two years. The gestation period is about 108 days, and the litter size varies from one to six cubs, with two to four on average. The newborn cub is helpless and blind and has a thick, dark-spotted coat, the markings of which usually disappear with maturity. Cubs are able to follow their mothers at around 3 months of age and are weaned by 6 or 7 months. They begin participating in kills by 11 months but probably cannot survive on their own until they are 2 years old. They reach sexual maturity at three or four years of age. There is a high mortality rate among cubs, and wild adults seldom live more than 8 or 10 years, chiefly owing to attacks by humans, other lions, or the effects of kicks and gorings from intended prey animals. Lions have a potential longevity of thirty years, but they seldom reach this age in the wilds.


Ancestors

The cat's early ancestors were primitive carnivores called creodonts. From the creodonts evolved a group of forest-dwelling hunters known as the miacid carnivores. The miacids were small; they had long bodies and tails and short, flexible limbs. The cats, civets, weasels, dogs and probably the seals and walruses as well are direct descendants of the miacids.

The first cats appeared about 35.000.000 years ago. These animals developed formidable upper fangs, up to 8 inches in length - which accounts for their popular name of sabre-toothed tigers. Sabre-tooths of various kinds roamed the forests and plains of all continents except Australia; they died out when the thick-skinned mammoths and other herbivores on which they preyed became scarce.

The modern felines probably evolved from early sabre-toothed cats, whose canine teeth where smaller than those of the most sabre-tooths. While the sabre-toothed cats were specialised for killing large, heavy and slow-moving herbivores, the true felines became adapted for hunting and catching agile animals, both large and small. Today, felines are among the most widespread and successful of mammalian groups. They inhabit all major land areas, with the exception of Australia, Madagascar and the oceanic islands. They are, however, most abundant in tropical regions.

During the late Pleistocene Epoch (1,600,000 to 10,000 years ago), lions had an extremely wide geographic distribution and ranged over all of North America and Africa, most of the Balkans, and across Anatolia and the Middle East into India. They disappeared from North America about 10,000 years ago, from the Balkans about 2,000 years ago, and from Palestine during the Crusades. Once lions were common in Eastern Europe and the Middle East but were hunted to extinction. The wanton slaughter of the lion (or any animal for that matter) is unforgivable. By the late 20th century their numbers had dwindled to a few tens of thousands, and those outside of national parks were rapidly losing their habitat to agriculture. Their future protection in Tanzania's Serengeti and other national parks seemed secure, however, partly because of the animals' tourist appeal.


Taxonomic hierarchy

Each of the individual animals on this site have a genus and species name, such as Panthera leo, beneath their common name. These are the two lowest levels of the classification system. In it's most basic form, the system actually has seven levels. For example, the full classification of an African lion would be as follows:

LEVEL CLASSIFICATION
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Family: Felidae
Genus: Panthera
Species: Leo

Man-eating

Sometimes a lion finds man's domestic animals easier to catch than its natural prey; rarely it becomes a man-eater. But it can happen that a lion, and tiger, too, develop a taste for human flesh because of various reasons. There is a possibility that certain individuals lose their innate fear of man by killing a person in a chance attack. Once successful in man-killing, these cats become less cautious toward man and find him an easier prey than the hoofed animals on which they usually feed. A period of famine, too, may embolden a big cat and make it a man-eater. This is especially true of a female nursing young; it may account for the fact that most man-eaters are females. In the Ruindi-Rutshuru plain, in Central Africa, man-eating reached epidemic proportions among lions around 1860 and again between 1904-09. This behaviour was probably caused by an outbreak of a fatal disease among topis and kobs, antelopes common to the area, that decreased their numbers dramatically.

A lion will not ordinarily attack a human being unless suddenly startled, bothered, wounded or diseased. Aged individuals are known to stalk and kill a person, but usually old ones exist on scorpions, insects and small rodents.

There was a similar situation in East Africa at the end of the 19th century. The notorious man-eating lions of Tsavo were two maneless male lions that killed 141 railroad workers in Kenya in 1898. Lt. Col. J.H. Patterson was the man who finally killed the man-eating lions of Tsavo; he recounts his experience in his booklet "The man-eating lions from Tsavo" he wrote in 1925. Includes black and white photographs.


Things worth knowing

In captivity, the lion has been induced to mate with other big cats. The offspring of a lion and a tigress is called a liger; that of a tiger and a lioness, a tigon; that of a leopard and a lioness, a leopon.

The Philadelphia Zoo was the first in North America to display and breed the White Lion. The White Lion is not an albino. It is the product of recessive genes.